Fig. 1

Two male nuptial traits in Leptobrachium boringii, maxillary spines and loose skin, and ten sampling locations of skin sections. 1: dorsal head; 2: back; 3: dorsal forearm; 4: dorsal distal hindlimb; 5: lateral abdomen; 6: ventral head; 7: chest; 8: posterior belly; 9: ventral forearm; 10: ventral proximal hindlimb